Curriculum Vitae

BS, education/zoology, Truman State University (formerly N.E. Missouri State University), 1968; MS, entomology, Utah State University, 1971; PhD, entomology, Utah State University, 1975 Retired former owner of Whitworth Pest Solutions, Inc. Washington State University Adjunct Professor in Entomology since 2006.

Blow Fly Publications

Bennett, G. F. & Whitworth, T. L. (1991)  Studies on the life-history of some species of Protocalliphora (Diptera, Calliphoridae). Canadian Journal of Zoology, 69, 2048-2058.

Bennett, G. F. & Whitworth, T. L. (1992) Host, nest, and ecological relationships of species of Protocalliphora (Diptera, Calliphoridae). Canadian Journal of Zoology, 70, 51-61.

Capelle, K. J. & Whitworth, T. L. (1973) The distribution and avian hosts of Carnus hemapterus (Diptera: Calliphoridae) in North America. Journal of Medical Entomology, 10, 525-526.

Daoust, S. P., Savage, J., Whitworth, T. L., Be'Lisle, M., Brodeur, J. (2012) Diversity and abundance of ectoparasitic blow flies Protocalliphora (Diptera: Calliphoridae) and their Nasonia (Hymenoptera: Pteromalidae) parasitoids in Tree Swallow nests within agricultural lands of southern Quebec, Canada. Annals of the Entomological Society of America, 105, 471-478.

Dawson, R. D., Hillen, K. K. & Whitworth, T. L. (2005) Effects of experimental variation in temperature on larval densities of parasitic Protocalliphora (Diptera: Calliphoridae) in nests of tree swallows (Passeriformes: Hirundinidae). Environmental Entomology, 34, 563-568.

Dawson, R. D., Whitworth, T. L. & Bortolotti, G. R. (1999) Bird blow flies, Protocalliphora (Diptera: Calliphoridae), in cavity nests of birds in the boreal forest of Saskatchewan. Canadian Field-Naturalist, 113, 503-506.

Gentes, M. L., Whitworth, T. L., Waldner, C., Fenton, H. & Smits, J. E. (2007) Tree swallows (Tachycineta bicolor) nesting on wetlands impacted by oil sands mining are highly parasitized by the bird blow fly Protocalliphora spp. Journal of Wildlife Diseases, 43, 167-178.

Jewiss-Gaines, A., Marshall, S. A. & Whitworth, T. L. (2012) Cluster flies (Calliphoridae: Polleniinae: Pollenia) of North America. Canadian Journal of Arthropod Identification, 19, 1-22.

Jones, N., Whitworth, T. L. & Marshall, S. A. (2019) Blow flies of North America: Keys to the subfamilies and genera of Calliphoridae, and to the species of the subfamilies Calliphorinae, Luciliinae and Chrysomyinae. Canadian Journal of Arthropod Identification, 1-191. http://dx.doi.org/10.3752/cjai.2019.39

Kozlov, M. V. & Whitworth, T. L., (2002) Populations densities and diversity of Calliphoridae (Diptera) around a nickel-copper smelter at Monchegorsk, northwest Russia Entomologica Fennica, 13, 98-104.

Marshall, S. A., Whitworth, T. L. & Roscoe, L. (2011) Blow flies (Diptera: Calliphoridae) of eastern Canada with a key to Calliphoridae subfamilies and genera of eastern North America, and a key to eastern Canadian species of Calliphorinae, Luciliinae, and Chrysomyinae. Canadian Journal of Arthropod Identification, 1-93.

Revels, M. R. & Whitworth, T. L. (2005) First record of Swainson's Warbler parasitism by Protocalliphora blow fly larvae. Wilson Bulletin 117, 199-200.

Rowland, S. M., Whitworth, T. L., Jones, M., Dooley, J., Chameroy, E. & Gordon, T. (2023) The bison and the blow fly: using puparia of the black blow fly (Phormia regina: Diptera, Calliphoridae) to constrain the season of death and taphonomic history of an early historic-age bison, Carson city, Nevada, USA. Palaios, 38, 22-30. Click Here To Read The Article

Sabrosky, C. W., Bennett, G. F. & Whitworth, T. L. (1989) Bird Blow Flies (Protocalliphora) in North America (Diptera: Calliphoridae) with notes on Palearctic species. Washington, D C: Smithsonian Institute Press, p. 312.

Sanford, M. R., Whitworth, T. L. & Phatak, D. R. (2014) Human Wound Colonization by Lucilia eximia and Chrysomya rufifacies (Diptera: Calliphoridae): Myiasis, Perimortem, or Postmortem Colonization? Journal of Medical Entomology, 51, 716-719.

Tantawi, T. I. & Whitworth, T. L. (2014) First record of Lucilia bufonivora Moniez, 1876 (Diptera: Calliphoridae) from North America and key to North American species of the L. bufonivora species group. Zootaxa, 3881, 101-124.

Whitworth, T. L. (1971) A Study of the biology of the species of Protocalliphora in the northern Wasatch Range, unpublished MS Thesis. Utah State University, p. 58.

Whitworth, T. L. (1976) Host and habitat preferences, life history, pathogenicity and population regulation in species of Protocalliphora Hough (Diptera: Calliphoridae). PhD disseration. Utah State University, Dissertation Abstracts International, published 1977, p. 144.

Whitworth, T. L. (2002)  Two new species of North American Protocalliphora Hough (Diptera: Calliphoridae) from bird nests. Proceedings of the Entomological Society of Washington, 104, 801-811.

Whitworth, T. L. (2003)  A key to the puparia of 27 species of North American Protocalliphora Hough (Diptera: Calliphoridae) from bird nests and two new puparial descriptions. Proceedings of the Entomological Society of Washington, 105, 995-1033.

Whitworth, T. L. (2003) A new species of North American Protocalliphora Hough (Diptera: Calliphoridae) from bird nests. Proceedings of the Entomological Society of Washington, 105, 664-673.

Whitworth, T. L. (2006) Keys to the genera and species of blow flies (Diptera: Calliphoridae) of America north of Mexico. Proceedings of the Entomological Society of Washington, 108, 689-725. Click here for Spanish language version.
Tips For Using The Keys More Effectively And Errata
Tips For Using The Keys More Effectively And Errata (Spanish language version)

Whitworth, T. L. (2010) Keys to the blow fly species (Diptera Calliphoridae) of America, north of Mexico. In: J. H. Byrd & J. L. Castner (Eds), Forensic Entomology, the utility of arthropods in legal investigations. CRC Press, Boca Raton, Chapter 19, 581-625.

Whitworth, T. L. (2010) Keys to the genera and species of blow flies (Diptera: Calliphoridae) of the West Indies and description of a new species of Lucilia Robineau-Desvoidy. Zootaxa, 2663, 1-35. Available online at http://www.mapress.com/zootaxa/list/2010/2663.html

Whitworth, T. L. (2012) Identification of Neotropical blow flies of the genus Calliphora Robineau-Desvoidy (Diptera: Calliphoridae) with the description of a new species. Zootaxa, 3209, 1-27. Available online at http://www.mapress.com/zootaxa/2012/f/zt03209p027.pdf

Whitworth, T. L. (2014) A revision of the Neotropical species of Lucilia Robineau-Desvoidy (Diptera: Calliphoridae). Zootaxa, 3810, 1-76. Available online at http://www.mapress.com/zootaxa/2014/f/zt03810p076.pdf 

Whitworth, T. L. & Rognes, K. (2014) Musca purpurascens Walker, 1836 (Insecta, Diptera, Calliphoridae): proposed conservation of prevailing usage of name by setting aside the unidentifiable female holotype and replacing it with a male neotype. Bulletin of Zoological Nomenclature, 71, 166-169. Available online at this link.

Whitworth, T. L. & Bennett, G. F. (1992) Pathogenicity of larval Protocalliphora (Diptera: Calliphoridae) parasitizing nestling birds. Canadian Journal of Zoology, 70, 2184-2191.

Whitworth, T. L., Dawson, R. D., Magalon, H. & Baudry, E. (2007) DNA barcoding cannot reliably identify species of the blowfly genus Protocalliphora (Diptera : Calliphoridae). Proceedings of the Royal Society B-Biological Sciences.

Whitworth, T. L., Bolek, M. G. & Arias-Robledo, G. (2020) Lucilia bufonivora, not Lucilia silvarum (Diptera: Calliphoridae), causes myiasis in anurans in North America with notes about Lucilia elongata and Lucilia thatuna. Journal of Medical Entomology, 58, 88-92. https://doi.org/10.1093/jme/tjaa143

Whitworth, T. L. (2020) Keys to the blow fly species (Diptera Calliphoridae) of America, north of Mexico. In: J. H. Byrd & J. K. Tomberlin (Eds), Forensic Entomology, the utility of arthropods in legal investigations, third edition. CRC Press, Boca Raton, FL, pp. 413-443.

Whitworth, T. L. & Amat, E. (2023) A new species of Lucilia Robineau-Desvoidy (Diptera, Calliphoridae) from the Amazon region and its potential distribution areas. Zootaxa, 5270, 133-138. Click Here To Read The Article